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Mr. Larijani, the top national security official and a confidante of the former supreme leader, expertly navigated Iran's internal politics and led the brutal crackdown on protests this year.
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The killing, which Iran confirmed, could embolden Iranian hard-liners, as the U.S.-Israeli bombing campaign continues.
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Ali Larijani, Iran's lead security official, and Basij commander Gholamreza Soleimani were "eliminated," Israel said. Iran has not commented on the strikes.
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Larijani has long been seen as one of the Islamic Republic's most experienced and influential policy makers.
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